Diversity Over Space and Time

Latitudinal distribution of surface area

  • Surface area declines from equator to poles.
  • Distribution of land v. oceans has nearly reversed over geologic time.
  • Influences climatic gradients and area of different environments.

Diversity Gradients Have Three Major Influences

  • Ecological interactions and tolerances to environment: species inhabit areas with appropriate resources and climate.
  • Evolutionary processes: speciation and extinction rates vary over time and space.
  • Earth history, including plate motions, climate change, and landscape change, determines the geography of environmental gradients and barriers to geographic-range shifts.

Earth History and Diversity Gradients

  • We observe high diversity in tropical latitudes today, and the climatic tropics are restricted to low latitudes.
  • 50 million years ago, climatically tropical environments extended over much of the earth, encompassing a much larger area of land and sea than today.
  • Global cooling caused tropical ecosystems to shrink to low latitudes, temperate ecosystems to expand, and boreal ecosystems to appear.
  • Modern tropical diversity reflects the legacy of a long existence and large area of this ecosystem.
  • High-latitude boreal forest and tundra ecosystems are geologically young.

Summary

  • Species diversity displays strong geographic gradients, especially the increase in species diversity at low latitudes and in regions of topographic complexity.
  • These gradients in species diversity follow latitudinal temperature gradients today.
  • Over geologic time, these gradients were much shallower, corresponding to shallow temperature gradients.
  • Both the age and the historically large area of tropical ecosystems contribute to high tropical diversity today.