indicates student or postdoc advisee


Faith JT, Braun DR, Davies B, DeSantis LRG, Douglass MJ, Esteban I, Hare V, Levin NE, Luyt J, Pickering R, Power MJ, Sealy J, Deano S. 2020. Ecometrics and the paleoecological implications of Pleistocene faunas from the western coastal plains of the Cape Floristic Region, South Africa, Journal of Quaternary Science, doi:10.1002/jqs.3247.

Frost SR, Simpson SW, Levin NE, Quade J, Rogers MJ, Semaw S. 2020. Fossil Cercopithecidae from the early Pliocene Sagantole Formation at Gona, Ethiopia. Journal of Human Evolution, 144: 102789.

Semaw S, Rogers MJ, Simpson, SW, Levin NE, Quade J, Dunbar N, McIntosh WC, Cáceres I, Stinchcomb GE, Holloway RL, Brown FH, Butler RF, Stout D, Everett M. 2020. Co-occurrence of Acheulian and Oldowan artifacts with Homo erectus cranial fossils from Gona, Afar, Ethiopia. Science Advances, Vol. 6, no. 10, eaaw4694. link to article. Press: UM news release.

*Bedaso ZK, *DeLuca NM, Levin NE, Zaitchik BF, Waugh DW, Wu S-Y, Harman CJ, Shanko D, 2020. Spatial and temporal variation in the isotopic composition of Ethiopian precipitation. Journal of Hydrology,, link to article.

Levin NE, Simpson SW, Quade J, Everett ME, Frost SR, Rogers MJ, Semaw S. The 6-million-year record of ecological and environmental change at Gona, Afar Region, Ethiopia, African Paleoecology and Human Evolution, Cambridge University Press, SC Reynolds and R Bobe (editors).


Saylor BZ, Gibert L, Deino A., Alene, M, Levin NE, Melillo SM, Peaple MD, Feakins SJ, Bourel B, Barboni D, Novello A, Sylvestre F, Mertzman SA, Haile-Selassie Y, 2019. Age and context of mid-Pliocene hominin cranium from Woranso-Mille, Ethiopia. Nature 573, 220-224. link to article. Press: UM news release, CNN article, CMNH announcement with video.

Simpson SW, Levin NE, Quade J, Rogers MJ, Semaw S. 2019. Ardipithecus ramidus postcranial fossils from the Gona Project Area, Afar Regional State, Ethiopia. Journal of Human Evolution, 129: 1-45. link to article


Potts R, Behrensmeyer AK, Faith JT, Tryon CA, Brooks AS, Yellen JE, Deino AL, Kinyanjui R, Clark JB, Haradon C, Levin NE, Meijer HJM, Veatch EG, Owen RB, Renaut RW. 2018. Environmental dynamics during the onset of the Middle Stone Age in eastern Africa. Science 360 (6384): 86-90. doi: 10.1126/science.aao2200 link to article

*Lehmann SB, Levin NE, Braun DR, Stynder DD, Zhu M, le Roux P, Sealy J.  Environmental and ecological implications of strontium isotope ratios in mid-Pleistocene fossil teeth from Elandsfontein, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology, 490: 84-94. doi: 10.1016/j.palaeo.2017.10.008 link to article

Forrest F, Stynder DD, Bishop LC, Levin NE, *Lehmann SB, Patterson DB, Matthews T, Braun DR. Zooarchaeological reconstruction of newly excavated Middle Pleistocene deposits from Elandsfontein, South Africa. Journal of Archaeological Science: Reports, 17: 19-29. doi: 10.1016/j.jasrep.2017.10.034 link to article


Blumenthal SA, Levin NE, Brown FH, Brugal J-P, Chritz KL, Harris JM, Jehle GE, Cerling TC. 2017. Aridity and hominin environments. Proceedings of the National Academy of Sciences USA, 114 (28): 7331-7336. doi: 10.1073/pnas.1700597114 link to article

*Li S, Levin NE, Soderberg K, Dennis KJ, Caylor KK. 2017. Triple oxygen isotope composition of leaf waters in Mpala, centra Kenya, Earth and Planetary Science Letters, 468: 38-50. doi: 10.1016/j.epsl.2017.02.015 link to article 


Haile-Selassie Y, Melillo SM, Ryan TM, Levin NE, Saylor BZ, Deino A, Mundil R, Scott G, Alene M, Gibert L. 2016. Dentognathic remains of Australopithecus afarensis from Nefuraytu (Woranso-Mille, Ethiopia): comparative description, geology, and paleoecological context. Journal of Human Evolution 100: 35-53. doi: 10.1016/j.jhevol.2016.08.003. link to article

*Lehmann SB, Braun DR, Dennis KJ, Patterson DB, Stynder DD, Bishop LC, Forrest F, Levin NE. 2016. Stable isotopic composition of fossil mammal teeth and environmental change in southwestern South Africa during the Pliocene and Pleistocene, Palaeogeography, Palaeoclimatology, Palaeoecology, 457: 396-408. doi: 10.1016/j.palaeo.2016.04.042 link to article

Patterson DB, *Lehmann SB, Matthews T, Levin NE, Stynder D, Braun DR. 2016. Stable isotope ecology of Cape dune mole-rats (Bathyergus suillus) from Elandsfontein, South Africa: implications for C4 vegetation and hominin paleobiology in the Cape Floral Region, Palaeogeography, Palaeoclimatology, Palaeoecology, 457: 409-421. doi: 10.1016/j.palaeo.2016.04.044 link to article


Levin NE, Haile-Selassie Y, Frost SR and Saylor BZ. 2015. Dietary change among hominins and cercopithecids in Ethiopia during the early Pliocene. Proceedings of the National Academy of Sciences USA, 117: 12304-12309, doi: 10.1073/pnas.1424982112. link to article

Cerling TE, Andanje SA, Blumenthal SA, Brown FH, Chritz KL, Harris JM, Hart JA, Kirera FM, Kaleme P, Leakey LN, Leakey MG, Levin NE, Manthi FK, Passey BH and Uno KT. 2015. Dietary changes of large herbivores in the Turkana Basin, Kenya from 4 to 1 Ma. Proceedings of the National Academy of Sciences USA, 112: 11467-11472, doi:10.1073/pnas.1513075112. link to article

Levin NE. 2015. Environment and climate of early human evolution. Annual Review of Earth and Planetary Sciences, 43: 405-429. link to article

*Li S, Levin NE, and Chesson LA. 2015. Continental scale variation in 17O-excess of meteoric waters in the United States. Geochimica et Cosmochimica Acta, 164: 110-126. link to article

Haile-Selassie Y, Gibert L., Melillo SM, Ryan TM, Alene, M, Deino A, Levin NE, Scott, G and Saylor BZ. 2015. New species from Ethiopia further expands Middle Pliocene hominin diversity. Nature, 521: 483-488. doi:10.1038/nature14448. link to article

Simpson SW, Kleinsasser L, Quade J, Levin NE, McIntosh WC, Dunbar N, Semaw S, Rogers M. 2015. Late Miocene hominin teeth from the Gona Paleoanthropological Research Project area, Afar, Ethiopia. Journal of Human Evolution, 81: 68-82. link to article


Passey BH, Hu H, Ji H, Montanari S, Li S*, Henkes GA, Levin NE. 2014. Triple oxygen isotopes in biogenic and sedimentary carbonates. Geochimica et Cosmochimica Acta, 141: 1-25. link to article

Levin NE, Raub TD, Dauphas N, Eiler JM. 2014. Triple oxygen isotope variations in sedimentary rocks. Geochimica et Cosmochimica Acta, 139: 173-189. PDF DATA

Eiler JM, Bergquist B, Bourg I, Cartigny P, Farquhar J, Gagnon A, Guo W, Halevy I, Hofmann A, Levin N, Schauble E, Stolper D. 2014. Frontiers of stable isotope geoscience. Chemical Geology, 372: 119-143. PDF

Simpson SW, Quade J, Levin NE, Semaw S. 2014. The female Homo erectus pelvis from Gona: Response to Ruff. 2010. Journal of Human Evolution, 68: 32-35. PDF


Braun DR, Levin NE, Stynder D, Herries AIR, Archer W, Forrest F, Roberts DL, Bishop LC, Matthews T, Lehmann SB*, Pickering R, Fitzsimmons K. 2013. Mid-Pleistocene Hominin Occupation at Elandsfontein, Western Cape, South Africa. Quaternary Science Reviews, 82: 145-166. PDF

Berman ESF, Levin NE, Landais A, Li S*, Owano T. 2013. Measurement of δ18O, δ17O, and 17O-excess in water by off-axis integrated cavity output spectroscopy and isotope ratios mass spectrometry. Analytical Chemistry, 85(21): 10392-10398. doi: 10.1021/ac402366t. PDF

Braun DR, Levin NE, Roberts D, Stynder D, Forrest FF, Herries AI, Matthews T, Bishop L, Archer W, Pickering R. 2013. Initial investigations of Acheulean hominin behaviour at Elandsfontein. In: The Archaeology of the West Coast of South Africa (eds. A Jerardino, A Malan, D Braun). Archaeopress, Oxford, pp. 10-23. PDF

Feakins, SJ, Levin NE, Liddy HM, Sieracki A, Eglinton TI, and Bonnefille R. 2013. Northeast African vegetation change over 12 m.y. Geology, 41: 295-298, doi:10.1130/G33845.1. PDF

Quade J and Levin NE. 2013. East African hominid paleoecology: isotopic evidence from paleosols. In: Early Hominin Paleoecology (eds. M Sponheimer, J Lee-Thorp, K Reed, P Ungar). University of Colorado Press, pp. 59-102, doi: 10.5876/9781607322252:C03.

Zaitchik BF and Levin NE. 2013. Understanding the Dynamics of the Tropical African Climate. Eos 94 (23): 209.


Haile-Selassie Y, Saylor BZ, Deino A, Levin NE, Alene M, and Latimer BM. 2012. A new hominin foot from Ethiopia shows multiple Pliocene bipedal adaptations, Nature, 483: 565-569, doi:10.1038/nature10922. PDF


Cerling, TE, Levin NE, and Passey BH. 2011. Stable Isotope Ecology in the Omo-Turkana Basin, Evolutionary Anthropology, 20(6): 228-237, doi: 10.1002/evan.20326. PDF

Cerling TE, Wynn JG, Andanje SA, Bird MI, Korir DK, Levin NE, Mace W, Macharia AN, Quade J, and Remien CH. 2011. Woody cover and hominin environments in the past 6 million years, Nature, 476: 51-56, doi:10.1038/nature10306. PDF

Levin NE, Brown FH, Behrensmeyer AK, Bobe R, and Cerling TE. 2011. Paleosol carbonates from the Omo Group: isotopic records of local and regional environmental change in East Africa. Palaeogeography, Palaeoclimatology, Palaeoecology, 307:75-89, doi: 10.1016/j.palaeo.2011.04.026. PDF DATA


Passey BH, Levin NE, Cerling TE, Brown FH, and Eiler JM. 2010. High-temperature environments of human evolution in East Africa based on bond ordering in paleosol carbonates. Proceedings of the National Academy of Sciences USA, 107: 11245-11249, doi:10.1073/pnas.1001824107. PDF

Braun DR, Harris JWK, Levin NE, McCoy JT, Herries AIR, Bamford MK, Bishop LC, Richmond BG, Kibunjia M. 2010. Early hominin diet included diverse terrestrial and aquatic animals 1.95 Ma in East Turkana, Kenya. Proceedings of the National Academy of Sciences USA, 107: 10002-10007,doi:10.1073/pnas.1002181107. PDF

Cerling TE, Levin NE, Quade J, Wynn JG, Fox DL, Kingston JD, Klein RG, and Brown FH. 2010. Comment on the Paleoenvironment of Ardipithecus ramidus: Science, 328: 1105-d. PDF


Levin NE, Zipser EJ, and Cerling TE, 2009, Isotopic composition of waters from Ethiopia and Kenya: Insights into moisture sources for eastern Africa. Journal of Geophysical Research, D23306; doi: 10.1029/2009JD012166. PDF DATA


Levin NE, Simpson SW, Quade J, Cerling TE, and Frost SR, 2008, Herbivore enamel carbon isotopic composition and the environmental context of Ardipithecus at Gona, Ethiopia, in Quade J, and Wynn JG, eds., The Geology of Early Humans in the Horn of Africa: Geological Society of America Special Paper 446, p. 215–234, doi: 10.1130/2008.2446(10). PDF DATA

Quade J, Levin NE, Simpson SW, Butler R, McIntosh WC, Semaw S, Kleinsasser L, Dupont-Nivet G, Renne P, Dunbar N, 2008, The Geology of Gona, Ethiopia, in Quade J, and Wynn JG, eds., The Geology of Early Humans in the Horn of Africa: Geological Society of America Special Paper 446, p. 1-31, doi: 10.1130/2008.2446(01). PDF

Kleinsasser LL, Quade J, McIntosh WC, Levin NE, Simpson SW, Semaw S. Stratigraphy and geochronology of the late Miocene Adu-Asa Formation at Gona, Ethiopia, in Quade J, and Wynn JG, eds., The Geology of Early Humans in the Horn of Africa: Geological Society of America Special Paper, 446, p.33-65, doi: 10.1130/2008.2446(02). PDF

Simpson SW, Quade J, Levin NE, Butler R, Dupont-Nivet G, Everett M, Semaw S. 2008. A Female Homo erectus Pelvis from Gona, Ethiopia. Science, 322: 1089-1092. PDF

Cerling TE, Harris JM, Hart JA, Kaleme P, Klingel H, Leakey MG, Levin NE, Lewison RI, Passey BH. 2008. Stable Isotope Ecology of Hippopotamus amphibius in East Africa. Journal of Zoology, 276: 204-212, doi:10.1111/j.1469-7998.2008.00450.x. PDF


Passey BH, Cerling TE, Levin NE. 2007. Temperature dependence of oxygen isotope acid fractionation in modern and fossil tooth enamels. Rapid Communications in Mass Spectrometry, 21: 2853-2859. PDF

Levin NE, Cerling TE, Passey BH, Harris JM, Ehleringer JR. 2006. A stable isotope aridity index for terrestrial environments. Proceedings of the National Academy of Sciences, USA 103: 11201-11205. PDF DATA

Stout D, Quade J, Semaw S, Rogers MJ, Levin NE. 2005. Raw Material selectivity of the earliest stone toolmakers at Gona, Afar, Ethiopia. Journal of Human Evolution, 48: 365-380. PDF

Semaw S, Simpson SW, Quade J, Renne PR, Butler RF, McIntosh WC, Levin N, Dominguez-Rodrigo M, Rogers MJ. 2005. Early Pliocene hominids from Gona, Ethiopia. Nature, 433: 301-305. PDF

Levin NE, Quade J, Simpson SW, Semaw S, Rogers MJ. 2004. Isotopic evidence for Plio-Pleistocene environmental change at Gona, Ethiopia. Earth and Planetary Science Letters, 219: 93-110. PDF DATA

Quade J, Levin N, Semaw S, Stout D, Renne PR, Rogers MJ, Simpson SW. 2004. Paleoenvironments of the earliest stone toolmakers, Gona, Ethiopia. Geological Society of America Bulletin, 116 (11-12): 1529-1544. PDF

Data Compilations

Levin NE. 2013. Compilation of East Africa Soil Carbonate Stable Isotope Data. Integrated Earth Data Applications. doi: 10.1594/IEDA/100231.